Ascocarp Morphology. Ascocarp subglobose or lobed, orifice or cavity absent or present (rare), 1-10 cm in diameter, brown or black, with warts, warts angulated, color of spore-bearing tissue yellow or brown (hazel), veins narrow and numerous. Peridium 200-400 micron (Pegler et al. 1993), outermost layers 50-150 micron (Pegler et al. 1993), yellow-brown or red-brown, anatomical structure pseudoparenchymatic with angular cells, cell tangential dimension 10-15(-20) micron (Pegler et al. 1993), cell radial dimension 10-15(-20) micron (Pegler et al. 1993), innermost layers 150-300 micron, color hyaline, anatomical structure plectenchymatic, external cystidia absent. Sterile veins plectenchimatic.
Asci ans spores Morphology. Asci globose, major axis 60-90(-110) micron (Pegler et al. 1993), minor axis 55-70 micron (Pegler et al. 1993), number of spores per ascus 1-6(-7), irregularly clustered.
Spores ellipsoid or subglobose, major axis 25-36 micrometer lenght (Chevalier and Frochot, 1998), minor axis 18-28 micrometer lenght (Chevalier and Frochot, 1998), color yellow or pale brown or medium brown, reticulation; ornamentation length 1-2-3 micron; number of meshes along the major axis 3-5 (Pegler et al. 1993).
Distribution. Habitat temperate; Putative plant host Quercus, or Corylus, or Ostrya, or Carpinus, or Fagus, or Castanea, or Pinus, or Cedrus.
DNA polymorphism. ITS restriction pattern with EcoRI 400-450 bp; SphI less than 50 bp, or 50-100 bp, or 300-350 bp; HincII 400-450 bp; DNA sequence ID of the ITS1-5.8S-ITS2 regions U95175.
Mycorrhizae morphology. Mycorrhizae emanting hyphae absent, or rare; anatomy of emanating hyphae: clamps absent; color of emanating hyphae hyaline; color of very tip whitish, or ochre, yellowish brown; color of older parts ochre, yellowish brown, or brown; cystidia absent, or present; wolly; color ochre; intercalate swelling present; ramification absent; anatomy of the mantle pseudoparenchymatus with angular cells.